Lillie is a central character in the Pokémon Sun and Moon anime series, known for her gentle and caring nature. She has a strong bond with her Pokémon, Nebby, which she carries in her bag, playing a protective role throughout the series. Lillie's character development is significant as she overcomes her fear of touching Pokémon, which stems from a traumatic experience in her past. Her growth is closely tied to her relationship with her mother, Lusamine, and her brother, Gladion, as they navigate their roles within the Aether Foundation.